C
AFOLIICOLOUS LICHEN DISTRIBUTEDA
IN LICHENES FOLIICOLI EXSICCATI
C

 

Robert LÜCKING*

*Lehrstuhl für Pflanzensystematik, Universität Bayreuth, D-95447 Bayreuth, Germany

[Version: October 31st, 2000]

 

Since the early years of lichenology, there has been an enormous production of exsiccates [SAYRE, Mem. New York Bot. Garden 19: 1-174. (1969)]. Exsiccates serve for three different purposes: (1) Wide distribution of duplicates, instead of housing concentrated collections at one locality, to ensure their preservation; (2) establishment of reference collections; (3) presentation of taxonomic concepts by distribution of selected material. Early exsiccates which partly included foliicolous lichens were Edward Vainio: "Lichenes Brasilienses Exsiccati" (1891-1892) and Anonymous: "Kryptogamae Exsiccatae Editae a Museo Palatino Vindobonensi" (since 1894). In recent times, foliicolous lichens have been treated in Klaus Kalb: "Lichenes Neotropici" (since 1981), Antonín Vezda: "Lichenes Selecti Exsiccati" (1966-1991) and "Lichenes Rariori" (since 1992), and Harrie Sipman: "Lichenotheca Latinoamericana" (since 1990).

"Lichenes Foliicoli Exsiccati" is the first exsiccate exclusively dedicated to foliicolous lichens. It has been started to establish a reference set of foliicolous lichen taxa from all over the world, including regional and local variations of species, populations with particular characters, and distinctive ecological assemblies. The exsiccate comprises 25 numbers per fascicle and at least twelve duplicates, regularly distributed among the following institutions and private herbaria:

  • Botanischer Garten und Botanisches Museum Berlin-Dahlem (B)
  • Australian National Botanic Gardens, Canberra (CBG)
  • Institut für Botanik, Karl-Franzens-Universität Graz (GZU)
  • Klaus Kalb, Lichenologisches Institut Neumarkt (KALB)
  • Department de Botanique, Université de Liège (LG)
  • Botanische Staatssammlung München (M)
  • Staatliches Museum für Naturkunde, Stuttgart (STU)
  • Fytoteket, Department of Botany, University of Uppsala (UPS)
  • Institute of Ecology and Botany of the Hungarian Academy of Sciences, Vácrátót (VBI)
  • André Aptroot, Baarn (hb. Aptroot)
  • Thorsten Lumbsch, Universität Essen (hb. Lumbsch)
  • Antonín Vezda, Botanical Institute, Brno (hb. Vezda)

Each fascicle is compiled under a certain taxonomic, systematic, ecological, geographical, or historical aspect. Specimens distributed under a given number originate from the same phorophyte or phorophyte species, representing the same morphological state. The fascicles are accompanied by individual schedae which are each dedicated to a certain topic. The schedae are produced at a number of 100 per fascicle and can be ordered from the editor.

Ten fascicles have been issued so far, two in 1993, four in 1995, and four in 2000:

  • Fascicle I [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. I. Faculty of Natural Sciences (1993)]
    Tropical rain forest
  • Fascicle II [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. II. Faculty of Natural Sciences (1993)]
    Sieghard Winkler
  • Fascicle III [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. III. Faculty of Natural Sciences (1995)]
    Rolf Santesson
  • Fascicle IV [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. IV. Faculty of Natural Sciences (1995)]
    Rain forest canopy
  • Fascicle V [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. V. Faculty of Natural Sciences (1995)]
    Taxonomically difficult foliicolous lichens and concept of mixed collections
  • Fascicle VI [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. VI. Faculty of Natural Sciences (1995)]
    Island floras
  • Fascicle VII [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. VII. Faculty of Natural Sciences (2000)]
    Porina epiphylla group NEW!
  • Fascicle VIII [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. VII. Faculty of Natural Sciences (2000)]
    Ecuador NEW!
  • Fascicle IX [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. VII. Faculty of Natural Sciences (2000)]
    Tropical rain forest understory NEW!
  • Fascicle X [LÜCKING, Schedae ad Lichenes Foliicoli Exsiccati, Fasc. VII. Faculty of Natural Sciences (2000)]
    Antonín Vezda NEW!

Up to the present, 183 species in 52 genera have been distributed in "Lichenes Foliicoli Exsiccati". The specimens include 14 isotypes and five topotypes. Most species originate from the Neotropics, but due to the support of many colleagues, a number of paleotropical collections are currently prepared for further editions. The following colleagues are warmely thanked for their contributions: A. Aptroot, M. E. da S. Cáceres, E. Farkas, L. I. Ferraro, K. Kalb, the late H. MacKee, R. Santesson, Ch. Scheidegger, E. Sérusiaux, H. J. M. Sipman, H. Streimann, G. Thor, and A. Vezda.

 

FOLIICOLOUS LICHENS DISTRIBUTED IN LICHENES FOLIICOLI EXSICCATI
UP TO FASC. X (OCTOBER 2000):

 

FOLIICOLOUS LICHEN SPECIES ORIGIN Fasc. No.
AAA
Amazonomyces farkasiae (Lücking) Lücking, Sérus. & Thor Costa Rica II 33
AA[partly as Cryptothecia farkasiae Lücking] Costa Rica X 226
Amazonomyces sprucei (R. Sant.) Lücking, Sérus. & Thor Brazil III 72
AA[as Stirtonia sprucei R. Sant.]
Anisomeridium foliicola R. Sant. & Tibell Costa Rica III 51
Arthonia accolens Stirt. Costa Rica III 52
Costa Rica IX 201
Arthonia aciniformis Stirt. Costa Rica X 227
Arthonia cyanea Müll. Arg. Costa Rica III 53
Guyana X 228
Arthonia leptosperma (Müll. Arg.) R. Sant. Costa Rica III 54
Arthonia orbygniae (H. B. P. Upadhyay) Matzer TOPOTYPE Costa Rica I 3
AA[as Arthonia opegraphina Lücking]
Arthonia palmulacea (Müll. Arg.) R. Sant. Costa Rica IX 202
Arthonia trilocularis Müll. Arg. Costa Rica II 26
Aspidothelium fugiens (Müll. Arg.) R. Sant. Tanzania II 27
Ecuador VIII 179
Aspidothelium ornatum Lücking ISOTYPE Ecuador VIII 180
Aspidothelium scutellicarpum Lücking ISOTYPE Ecuador VIII 181
Asterothyrium anomalum var. pallidum Henssen & Lücking TOPOTYPE Costa Rica IV 76
AA[as Asterothyrium anomalum Kalb & Vezda]
Asterothyrium argenteum Müll. Arg. Costa Rica IV 77
Asterothyrium gyalideoides Henssen & Lücking ISOTYPE Costa Rica IV 78
Asterothyrium leucophthalmum (Müll. Arg.) R. Sant. Costa Rica IV 79
Asterothyrium pittieri Müll. Arg. Ecuador VIII 185
Asterothyrium rotuliforme (Müll. Arg.) Sérus. Costa Rica IV 80
Aulaxina minuta R. Sant. Costa Rica I 2
"Bacidia" brasiliensis (Müll. Arg.) Zahlbr. Guatemala I 4
"Bacidia" corallifera Lücking ISOTYPE Cocos Isl. VI 126
Ecuador VIII 196
"Bacidia" psychotriae (Müll. Arg.) Zahlbr. Brazil III 55
Bacidina apiahica (Müll. Arg.) Vezda Costa Rica III 56
Bacidina mirabilis (Vezda) Vezda Guyana IX 217
Bacidina pallidocarnea (Müll. Arg.) Vezda Costa Rica II 29
Badimia dimidiata (Babingt. ex Leight.) Vezda Costa Rica V 106
French Guiana VI 130
Badimia galbinea (Kremp.) Vezda Costa Rica I 5
Brazil III 57
Costa Rica V 107
Badimia montoyana Lücking ISOTYPE Cocos Isl. VI 129
Badimia pallidula (Kremp.) Vezda Costa Rica III 58
Badimia tuckermanii (R. Sant.) Lücking, Lumbsch & Elix New Caledonia VI 131
Bapalmuia lineata Lücking & Kalb Costa Rica IX 222
Bapalmuia palmularis (Müll. Arg.) Sérus. Costa Rica IX 221
Barubria fuscorubra (Vezda) Vezda Costa Rica X 229
Barubria flavescens Lücking ISOTYPE Costa Rica X 230
Bullatina aspidota (Vain.) Vezda & Poelt Costa Rica IV 81
Byssolecania fumosonigricans (Müll. Arg.) R. Sant. Costa Rica IX 224
Byssolecania hymenocarpa (Vain.) Kalb, Vezda & Lücking Costa Rica IX 225
Byssoloma amazonicum Kalb & Vezda Brazil III 59
Byssoloma chlorinum (Vain.) Zahlbr. Cocos Isl. VI 132
Costa Rica VI 133
Ecuador VIII 198
Byssoloma confusum Farkas & Vezda New Caledonia X 231
Byssoloma leucoblepharum (Nyl.) Vain. Canary Isl. II 30
Costa Rica IX 223
Byssoloma lueckingii Sérus. Costa Rica IV 82
Byssoloma ortizii Lücking Costa Rica V 102
Byssoloma subdiscordans (Nyl.) P. James Costa Rica IV 83
Byssoloma tricholomum (Mont.) Zahlbr. Costa Rica I 6
Guyana X 232
Byssoloma vezdanum Sérus. Guyana X 233
Byssoloma wettsteinii (Zahlbr.) Zahlbr. Cocos Isl. II 28
AA[partly as Bacidia psychotriae (Müll. Arg.) Zahlbr.] Costa Rica V 101
Cocos Isl. VI 127
Costa Rica VI 128
Calenia aggregata R. Sant. Costa Rica III 60
Calenia monospora Vezda Costa Rica IV 84
Calenia phyllogena (Müll. Arg.) R. Sant. Ecuador VIII 190
Calenia triseptata Zahlbr. Costa Rica I 7
AA[as Calenia submaculans R. Sant.]
Caleniopsis tetramera Lücking ISOTYPE Costa Rica X 234
Calopadia foliicola (Fée) Vezda Costa Rica IV 85
Brazil V 103a
Calopadia fusca (Müll. Arg.) Vezda Ecuador VIII 200b
Calopadia perpallida (Nyl.) Vezda Costa Rica II 31
Calopadia phyllogena (Müll. Arg.) Vezda Brazil V 103b
Guatemala X 235
Calopadia puiggarii (Müll. Arg.) Vezda Costa Rica IV 86
U.S.A. IV 104
Calopadia subcoerulescens (Zahlbr.) Vezda U.S.A. V 105
Chroodiscus australiensis Vezda & Lumbsch Costa Rica X 236
Chroodiscus coccineus (Leight.) Müll. Arg. Costa Rica I 8
Ecuador VIII 182
Coccocarpia domingensis Vain. Costa Rica II 32
Ecuador VIII 194
Coenogonium isidiiferum (Lücking) Lücking Costa Rica X 237
AA[as Dimerella isidiifera Lücking]
Coenogonium interpositum Nyl. Ecuador VIII 191
Coenogonium queenslandicum (Kalb & Vezda) Lücking Australia IX 215
AA[as Dimerella queenslandica Kalb & Vezda]
Cryptothecia candida (Kremp.) R. Sant. Costa Rica I 9
Cryptothecia inexspectata Thor Papua New Guinea IX 203
Coenogonium hypophyllum (Vezda) Kalb & Lücking French Guiana III 61
AA[partly as Dimerella hypophylla Vezda] Ecuador VIII 192
Costa Rica IX 216
Coenogonium subluteum (Rehm) Kalb & Lücking Cocos Isl. VI 134
AA[as Dimerella epiphylla (Müll. Arg.) Malme]
"Dimerella" vezdana Lücking Ecuador VIII 193
Echinoplaca atrofusca R. Sant. Costa Rica IV 87
Ecuador VIII 188
Echinoplaca epiphylla Fée Costa Rica I 10
Echinoplaca furcata Sérus. Ecuador VIII 189
Echinoplaca hymenocarpoides (Vain.) Lücking Costa Rica IV 88
Echinoplaca lucernifera Kalb & Vezda Costa Rica X 238
Echinoplaca pellicula (Müll. Arg.) R. Sant. Guyana II 34
Echinoplaca serusiauxii Lücking Ecuador VIII 187
Echinoplaca verrucifera Lücking Costa Rica X 239
Enterographa angustissima (Vain.) R. Sant. Costa Rica I 11
Enterographa byssoidea Lücking TOPOTYPE Costa Rica III 62
Fellhanera angustispora Lücking Costa Rica IX 220
Fellhanera avilezii Lücking ISOTYPE Cocos Isl. VI 135
Fellhanera bouteillei (Desm.) Vezda Costa Rica IV 89
Fellhanera bullata Kalb & Vezda New Caledonia VI 136
Fellhanera christiansenii Sérus. & Vezda Canary Isls. VI 137
Fellhanera fuscatula (Müll. Arg.) Vezda Costa Rica I 12
AA[partly as Fellhanera dominicana (Vain.) Vezda] Costa Rica IX 218
Fellhanera semecarpi (Vain.) Vezda Costa Rica II 35
Costa Rica IV 90
Fellhanera stanhopeae (Müll. Arg.) Lücking, Lumbsch & Elix Costa Rica III 63
Fellhanera subfuscatula Lücking Ecuador VIII 197
Fellhanera sublecanorina (Nyl.) Vezda Tanzania II 36
New Caledonia VI 138
Fellhanera substanhopeae Lücking Guyana X 240
Fellhanera verrucifera Lücking Costa Rica IX 219
Flavobathelium epiphyllum Lücking, Aptroot & Thor ISOTYPE Costa Rica X 241
Gyalectidium catenulatum (Cavalc. & A. A. Silva) Ferraro, Lücking & Sérus. Costa Rica III 64
AA[as Gyalectidium filicinum Müll. Arg.]
Gyalectidium filicinum Müll. Arg. Costa Rica I 13
Cocos Isl. VI 139
Gyalidea epiphylla Vezda Costa Rica I 14
Ecuador VIII 183
Gyalidea phyllophila Vezda Ivory Coast IX 212
Gyalideopsis verruculosa Vezda & Hafellner Australia II 37
Costa Rica X 242
Gyalideopsis vulgaris (Müll. Arg.) Lücking Costa Rica I 1
AA[as Actinoplaca vulgaris (Müll. Arg.) Vezda & Poelt]
Hippocrepidea nigra Sérus. ISOTYPE Papua New Guinea III 65
Lasioloma arachnoideum (Kremp.) R. Sant. Costa Rica IV 91
Cocos Isl. VI 140
Costa Rica VI 141
Loflammia epiphylla (Fée) Lücking & Vezda Costa Rica IV 94
AA[as Loflammia flammea (Müll. Arg.) Vezda]
Malcolmiella trailiana (Müll. Arg.) Vezda Brazil III 66
AA[as Lecidea trailiana Müll. Arg.]
Mazosia dispersa (Hedrick) R. Sant. Costa Rica III 67
Mazosia melanophthalma (Müll. Arg.) R. Sant. Australia II 38
Costa Rica IX 206
Mazosia phyllosema (Nyl.) Zahlbr. Costa Rica IX 205
Mazosia praemorsa (Stirt.) R. Sant. Costa Rica VI 142
Mazosia rotula (Mont.) A. Massal. Brazil II 39
Mazosia sorediifera Lücking & Matzer TOPOTYPE Costa Rica X 243
Mazosia tenuissima Lücking & Matzer Guyana IX 207
Mazosia tumidula (Stirt.) Müll. Arg. Costa Rica I 15
Ecuador VIII 176
French Guiana IX 208
Microtheliopsis uleana Müll. Arg. Costa Rica II 40
Microtheliopsis winkleri Lücking ISOTYPE Costa Rica II 41
Opegrapha filicina Mont. Costa Rica I 16
Opegrapha lambinonii Sérus. Costa Rica IX 204
Paratricharia paradoxa (Lücking) Lücking Costa Rica I 24
AA[partly as Tricharia paradoxa Lücking] Costa Rica IX 214
Phaeographis inconspicua (Fée) Müll. Arg. Costa Rica X 244
Phyllobathelium firmum (Stirt.) Vezda Costa Rica I 17
AA[partly as Phyllobathelium epiphyllum (Müll. Arg.) Müll. Arg.] Costa Rica X 245
Phyllophiale alba R. Sant. Costa Rica IV 95
Porina atriceps (Vain.) Vain. Costa Rica VII 151
Porina atropunctata Lücking & Vezda Costa Rica VII 152
Costa Rica VII 153
Costa Rica VII 154
French Guiana VII 155
Porina conica R. Sant. Vanuatu VII 157
Porina curtula Malme Guyana VII 175
Porina distans Vezda & Vivant Costa Rica III 68
Porina epiphylla (Fée) Fée Costa Rica VII 162
Costa Rica VII 163
Porina fusca Lücking Costa Rica II 43
Porina guianensis Lücking & Vezda Costa Rica VII 171
Porina impressa R. Sant. Australia X 246
Porina leptospermoides Müll. Arg. Costa Rica V 112
Costa Rica V 113
Costa Rica V 114
Porina limbulata (Kremp.) Vain. Costa Rica V 115
Costa Rica V 116
Costa Rica V 117
Australia V 118
Costa Rica V 123b
Porina lucida R. Sant. var. lucida Costa Rica VII 164
Cocos Isl. VII 165
Porina mazosioides Lücking & Vezda Madagascar VII 172
Porina minutissima Henssen, Lücking & Vezda Malaysia VII 161
Porina mirabilis Lücking & Vezda Costa Rica VII 166
Costa Rica VII 167
Costa Rica VII 168
Costa Rica VII 169
Porina nitidula Müll. Arg. Costa Rica IV 96
AA[as Pseudosagedia nitidula (Müll. Arg.) Hafellner & Kalb]
Porina papillifera (Stirt.) F. Schill. Costa Rica I 19
Ecuador VIII 177
Porina radiata Kalb, Lücking & Vezda Costa Rica VII 174
Porina repanda (Stirt.) Lücking & R. Sant. Costa Rica III 69
AA[partly as Pseudosagedia laticarpa (Lücking) Hafellner & Kalb] Ecuador VIII 178
Porina rubentior (Stirt.) Müll. Arg. Costa Rica V 119
Costa Rica V 120
Cocos Isl. VI 145
Costa Rica VI 146
Porina rufula (Kremp.) Vain. Costa Rica V 121
Costa Rica V 123a
Porina cf. rufula (Kremp.) Vain. French Guiana V 122
Porina similis Kalb & Vezda Australia VII 156
Porina sphaerocephaloides Farkas Madagascar VII 174
Porina subepiphylla Lücking & Vezda Costa Rica VII 158
Costa Rica VII 159
Ecuador VII 160
Porina vezdae Lücking Costa Rica X 247
Porina virescens (Kremp.) Müll. Arg. Vanuatu VII 170
Psorotheciopsis albomaculans (Rehm) R. Sant. Costa Rica IV 92
AA[as Linhartia albomaculans (Rehm) Sacc. & Syd.]
Psorotheciopsis guajalitensis Lücking ISOTYPE Ecuador VIII 184
Psorotheciopsis philippinensis (Rehm) Lücking Costa Rica IV 93
AA[as Linhartia vezdana Lücking] [Isotype!]
Scoliciosporum curvatum Sérus. ISOTYPE France II 44
Sporopodium flavescens (R. Sant.) Vezda Australia II 45
Sporopodium leprieurii Mont. Costa Rica III 71
Ecuador VIII 199
Sporopodium xantholeucum (Müll. Arg.) Zahlbr. Costa Rica I 21
Strigula antillarum (Fée) Müll. Arg. Costa Rica IV 97
Strigula concreta (Fée) R. Sant. Costa Rica V 124
Strigula janeirensis (Müll. Arg.) Lücking Costa Rica I 20
AA[partly as Raciborskiella janeirensis (Müll. Arg.) R. Sant.] Costa Rica IX 210
Strigula kaitokensis Sérus. & Polly ISOTYPE New Zealand VI 147
Strigula maculata (Cooke & Massee) R. Sant. Costa Rica I 22
Strigula multipunctata (G. Merr. ex R. Sant.) R. C. Harris Australia II 42
AA[as Phylloporis multipunctata (G. Merr. ex R. Sant.) Vezda]
Strigula nemathora Mont. Costa Rica II 46
Strigula nitidula Mont. Costa Rica IV 98
Costa Rica IV 99
Costa Rica V 125
Strigula obducta (Müll. Arg.) R. C. Harris Costa Rica V 108
AA[partly as Phylloporis obducta (Müll. Arg.) R. Sant. & Tibell] Ecuador IX 209
Strigula phyllogena (Müll. Arg.) R. C. Harris Costa Rica V 109
AA[partly as Phylloporis phyllogena (Müll. Arg.) Clem.] Costa Rica V 111a
Strigula platypoda (Müll. Arg.) R. C. Harris Costa Rica I 18
AA[partly as Phylloporis platypoda (Müll. Arg.) Vezda] Costa Rica V 110
Costa Rica V 111b
Cocos Isl. VI 143
Costa Rica VI 144
Strigula prasina Müll. Arg. Costa Rica III 70
AA[as Raciborskiella prasina (Müll. Arg.) R. Sant.]
Strigula schizospora R. Sant. Costa Rica II 47
Strigula smaragdula Fr. Cocos Isl. VI 148
Strigula subtilissima (Fée) Müll. Arg. Costa Rica II 48
Tapellaria bilimbioides R. Sant. Costa Rica I 23
Costa Rica VI 149
Tapellaria epiphylla (Müll. Arg.) R. Sant. Ecuador VIII 200a
Tapellaria nana (Fée) R. Sant. Costa Rica IV 100
Tapellaria puiggarii (Müll. Arg.) R. Sant. Guyana X 248
Tapellariopsis octomera Lücking Costa Rica X 249
Tricharia albostrigosa R. Sant. Costa Rica III 73
Tricharia farinosa R. Sant. Costa Rica II 49
Tricharia heterella (Stirt.) R. Sant. Costa Rica IX 213
Tricharia pseudosantessonii Lücking Costa Rica III 74
AA[partly as Tricharia santessonii D. Hawksw.] Ecuador VIII 186
Tricharia urceolata (Müll. Arg.) R. Sant. Costa Rica VI 150
Trichothelium annulatum (Karst.) R. Sant. Ecuador IX 211
Trichothelium bipindense F. Schill. Costa Rica III 75
Trichothelium epiphyllum Müll. Arg. Costa Rica I 25
Trichothelium longisporum Lücking TOPOTYPE Costa Rica II 50
Vezdaea foliicola Sérus. Ecuador VIII 195
Guadeloupe X 250